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Timdal, Einar
(2023).
Clades, clusters and species hypotheses in the crustose lichen genus Lecidea.
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Timdal, Einar
(2023).
Peltigera sect. Peltigera in Norway.
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Evankow, Ann; Bendiksby, Mika & Timdal, Einar
(2023).
Cryptic diversity of the lichen genus Psora in cryptogam collections .
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Timdal, Einar
(2022).
Diskusjon om lav tok 150 år.
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Bjorå, Charlotte Sletten; Bendiksby, Mika; Løfall, Bjørn Petter & Timdal, Einar
(2022).
Collections from the cold solving hot questions
.
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Timdal, Einar
(2022).
Lichens on heavy metal rock and soil in Norway.
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Timdal, Einar
(2022).
Nephroma helveticum-komplekset i Norden.
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Timdal, Einar
(2022).
Lav på tungmetall i Norge.
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Timdal, Einar
(2022).
Bruk av lav til farging – lav som egner seg til farging og hvorfor.
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Timdal, Einar
(2022).
Species hypotheses in Lecidea and Porpidia based on the DNA barcode marker.
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Evankow, Ann & Timdal, Einar
(2022).
Challenges & Unexpected Opportunities from constructing a Lichen DNA Barcode Database for Norway.
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Evankow, Ann & Timdal, Einar
(2022).
Searching for Psora in North America.
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Evankow, Ann; Bendiksby, Mika; Speed, James David Mervyn & Timdal, Einar
(2021).
INTEGRATIVE TAXONOMY OF THE Psoraceae.
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Evankow, Ann; Bendiksby, Mika & Timdal, Einar
(2020).
Integrative taxonomy of the Psoraceae (Lichenized Ascomycota).
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Lindblom, Louise; Blom, Hans H. & Timdal, Einar
(2020).
Xanthomendoza ulophyllodes – en allélav.
Lavbulletinen.
ISSN 1651-6435.
2020(1),
s. 32–38.
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Holien, Håkon & Timdal, Einar
(2018).
Tor Tønsberg: A birthday tribute.
Graphis Scripta.
ISSN 0901-7593.
30(6),
s. 30–33.
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Marthinsen, Gunnhild; Bendiksby, Mika; Brandrud, Tor Erik; Dima, Bálint; Hansen, Lars Ove & Johnsen, Arild
[Vis alle 8 forfattere av denne artikkelen]
(2017).
Discordance between morphology and DNA barcodes in lichens, fungi and insects.
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Dahl, Malin Stapnes; Bendiksby, Mika & Timdal, Einar
(2016).
Molecular systematics of the lichen genus Sporacestra and relatives (Ramalinaceae, Ascomycota).
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Toreskaas, Anne Karin; Bendiksby, Mika & Timdal, Einar
(2016).
Molecular systematics of the lichen genus Eschatogonia.
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Kistenich, Sonja Daniela; Timdal, Einar; Bendiksby, Mika & Ekman, Stefan
(2016).
Molecular phylogenetics and taxonomy of the lichen family Ramalinaceae.
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Dahl, Malin Stapnes; Bendiksby, Mika & Timdal, Einar
(2016).
Molecular systematics of the lichen genus Sporacestra and relatives (Ramalinaceae, Ascomycota).
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Toreskaas, Anne Karin; Bendiksby, Mika & Timdal, Einar
(2016).
Molecular systematics of the lichen genus Eschatogonia.
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Kistenich, Sonja Daniela; Timdal, Einar; Bendiksby, Mika & Ekman, Stefan
(2016).
Molecular phylogenetics and taxonomy of the lichen family Ramalinaceae.
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Haugan, Reidar; Timdal, Einar; Frisch, Andreas; Rui, Siri & Bendiksby, Mika
(2016).
Miriquidica – yet another non-monophyletic lichen genus rich in cryptic species.
Vis sammendrag
Background:
Miriquidica Hertel & Rambold comprises about 30 saxicolous, mainly alpine species worldwide. The genus was originally delimited by the presence of miriquidic acid in most of the species and by asci of the “Lecanora-type”. The species are often difficult to identify without chemical and microscopical investigations. Although they are common rock lichens in some regions in the Northern Hemisphere, many Miriquidica species remain poorly understood.
Material and Methods:
We have collected Miriquidica extensively throughout Fennoscandia, over wide climatic, geographical and ecological gradients. We studied this, and additional material of available Miriquidica species and presumed close relatives, using TLC, microscopy, and multi-locus DNA-sequencing. We have compiled a dataset with 187 accessions representing specimens collected as Miriquidica, their presumed close relatives, and appropriate outgroups. Some sequences were downloaded from genbank (mainly from Singh et al. 2013). The molecular work was undertaken as described in Bendiksby et al. (2013). A partitioned concatenated dataset of one mitochondrial (mtSSU) and two nuclear genetic regions (ITS and MCM7) was analysed phylogenetically using Maximum likelihood (ML) on the CIPRES Science Gateway (Miller et al. 2010). For the ML analysis, we used the RAxML-HPC black box with rapid bootstrapping and full ML analysis under the GTR+GAMMA approximation.
Results:
We have discovered several new species, many of them so-called cryptic and Miriquidica hence falls into the line of crustose genera showing a high degree of cryptic speciation (Fig. 1). Our results also show that Miriquidica, as currently circumscribed, is polyphyletic, i.e. one species (M. pycnocarpa) is extraneous in the genus. After its removal from Miriquidica, the genus remains paraphyletic, with species of Protoparmelia (cfr. Singh et al. 2013), Lecanora, Lecidea s.lat., and Myochroidea nested. We recognize several new species, of which we include two in Fig. 1 (M. scoriae and M. ymei), and recombine 11 others.
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Košuthová, Alica; Fačkovcová, Zuzana; Guttová, Anna; Holien, Håkon; Timdal, Einar & Wedin, Mats
[Vis alle 8 forfattere av denne artikkelen]
(2016).
DNA barcoding of lichens and biosystematics; the case studies of genera: Cladonia, Placynthium and Solenopsora.
Vis sammendrag
The main motivation for this project is a situation of shared research questions, complementary competences, and a desire for knowledge and data sharing, both taxonomically and methodologically, between Masaryk University and NTNU. On the lichenology side, there was a motivation for nourishing and fomalizing a dawning network of european lichen systematists. All involved, both the core participants (Bendiksby, Holien and Košuthová) and the accompanyiong scientists (Guttová, Steinová, Timdal, Wedin and Westberg), have some form of ongoing work with at least one participant of this project. However, their respective networks and competences are not the same, and new connections within the group has been made in the present project.
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Košuthová, Alica; Fačkovcová, Z.; Flatberg, Kjell Ivar; Guttová, Anna; Hájek, Michael & Hassel, Kristian
[Vis alle 16 forfattere av denne artikkelen]
(2015).
DNA barcoding of bryophytes and lichens, including biosystematic studies of selected groups.
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Bendiksby, Mika; Haugan, Reidar; Spribille, Toby & Timdal, Einar
(2015).
DNA barcoding and phylogenetics – An example from the Calvitimela aglaea lichen complex (Tephromelataceae, Lecanorales).
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Bendiksby, Mika & Timdal, Einar
(2014).
Molecular systematics and taxonomy of Southeast Asian members of the tropical rainforest genus Phyllopsora (Ramalinaceae).
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Olsen, Stein Alexander; Kumar, Rituvesh; Glenndal, Svein Gunnar; Larsson, Karl-Henrik; Prestø, Tommy & Timdal, Einar
[Vis alle 9 forfattere av denne artikkelen]
(2014).
MusitBotanikk - soppdatabasen, universitetsmuseenes samlingsdatabaser - versjon 1.2.
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Olsen, Stein Alexander; Kumar, Rituvesh; Glenndal, Svein Gunnar; Prestø, Tommy; Timdal, Einar & Rämä, Heini Emilia
[Vis alle 8 forfattere av denne artikkelen]
(2014).
MusitBotanikk - lavdatabasen, Universitetsmuseenes samlingsdatabaser versjon 1.2.
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Endresen, Dag Terje Filip; Svindseth, Christian; Timdal, Einar & Rindal, Eirik
(2014).
Crowdsourcing portal for transcription of label information for Norwegian natural history collections.
Vis sammendrag
The number of specimens in natural history museums and university-collections worldwide count approximately 1.2 to 3 billion specimens. These collections provide a unique resource for understanding biological and ecological processes across time and place. Norwegian natural history collections have been estimated to include a total of 8.5 million specimens. An estimated 65% of the specimens have been digitized (in some form) and 30% published to the GBIF portal. Recent large-scale digitization efforts in Norway have focused on the imaging of a large number of specimens accompanied by initial minimal database registration, limited to scientific name, year and the country where the specimen was collected. The registration of more complete label information is recognized to require laborious efforts and it was decided to explore a citizen science approach with a volunteer transcription portal. After exploring similar volunteer transcription portals (such as the portals from the Botanical Society of Britain & Ireland, Atlas of Living Australia, National Museum of Natural History in Paris and Smithsonian), the Natural History Museum in Oslo initiated contact and collaboration with the Notes from Nature team. A simplified software implementation, based on Notes from Nature, was developed at the Natural History Museum in Oslo during the first months of 2014 and launched on June 1st (http://gbif.no/transcribe/). The first collection of images to be made available for volunteer transcription included 2850 non-digitized specimens from the Lichen herbarium collected in Africa. Each image is registered three times by individual volunteers. Already during the first two weeks, more than 1000 of the specimens (40%) were completed. By early August, most of the (easy) labels were completed (70%). A new batch with images of non-digitized African Lichens, and also other collections such as the non-digitized specimens from the vascular plant herbarium, will soon be included to the portal.
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Haugan, Reidar; Bendiksby, Mika; Rui, Siri & Timdal, Einar
(2014).
Molecular phylogeny and taxonomy of the lichen genus Miriquidica (Ascomycota, Lecanoraceae).
Vis sammendrag
The lichen genus Miriquidica (Hertel & Rambold 1987) comprises about 30 saxicolous species worldwide. Its phylogeny remains unknown and several unknown/undescribed species may occur. Many Miriquidica species are difficult to identify without careful chemical (TLC) and microscope investigation. The genus was originally delimited by the presence of miriquidic acid (not all species) and anatomical characters of the apothecia, including asci of the “Lecanora-type”. Although Miriquidica species are not uncommon in alpine areas, and in some areas (e.g. in the mesotrophic Fennoscandian mountains) they represent a substantial part of the crustose lichen flora on rocks, the genus remains poorly represented in herbaria worldwide. We have collected Miriquidica species systematically throughout Norway, representing wide climatic, geographical and ecological gradients, and determined/studied this material through a combinations of different approaches (TLC, microscopy, DNA-sequencing). Our molecular phylogeny of the genus include multiple accessions of all available species as well as extra-Norwegian material. Most Miriquidica species will remain in a monophyletically circumscribed Miriquidica. Some Miriquidica species, however, are clearly extraneous in the genus, and a few species of Lecanora and Lecidea s.lat. belong in Miriquidica. Several new species are described and an extensive amount of phenotypically cryptic species have so far been discovered.
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Timdal, Einar
(2013).
Lichenized fungi.
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Timdal, Einar & Bendiksby, Mika
(2013).
Molecular phylogeny of Hypocenomyce - extreme polyphyly and morphological/ecological convergence.
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Timdal, Einar & Bendiksby, Mika
(2013).
Molecular phylogeny of Hypocenomyce - extreme polyphyly and morphological/ecological convergence.
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Bendiksby, Mika & Timdal, Einar
(2013).
Exploring species diversity and phylogeny of some obscure, pantropical rainforest lichens — DNA-barcoding, phylogenetics and taxonomy.
Vis sammendrag
Lichens occurring in the Tropics are generally poorly known, and crustose lichens in particular. We use DNA sequencing to test hypotheses about species delimitation and phylogeny of three obscure pantropical lichen genera, Eschatogonia, Krogia, and Phyllopsora. Although common epiphytes on rainforest trees, species delimitation and phylogeny of these genera are poorly understood; even their family belongings remain unsettled. Einar Timdal (NHM) is one of very few people with expert knowledge on these genera. He has published several revisions based on morphological and TLC studies of their secondary metabolites, but is still not able to determine more than about 90 % of the material and suspects there are much undiscovered and ‘hidden’ diversity. Only a single DNA sequence of Phyllopsora exists in GenBank (with obscure determination), and so far none of Eschatogonia and Krogia. An NHM-Småforsk grant has funded our molecular phylogenetic investigation of the smaller genera Eschatogonia and Krogia, as well as preliminary data for a Phyllopsora revision. About 70 Phyllopsora species are recognized today, of which 30 are described since the year 2000. Our molecular phylogenetic results and taxonomic conclusions on Eschatogonia and Krogia will be presented, as well as preliminary results on Phyllopsora.
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Bendiksby, Mika & Timdal, Einar
(2013).
Molecular systematics of lecideoid lichens – extreme polyphyly and a mess of cryptic species.
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Haugan, Reidar; Bendiksby, Mika; Rui, Siri & Timdal, Einar
(2013).
Molecular phylogeny and taxonomy of the lichen genus Miriquidica (Ascomycota, Lecanoraceae).
Vis sammendrag
The lichen genus Miriquidica (Hertel & Rambold 1987) comprises about 30 saxicolous species worldwide. Its phylogeny remains unknown and several unknown/undescribed species may occur. Many Miriquidica species are difficult to identify without careful chemical (TLC) and microscope investigation. The genus was originally delimited by the presence of miriquidic acid (not all species) and anatomical characters of the apothecia, including asci of the “Lecanora-type”. Although Miriquidica species are not uncommon in alpine areas, and in some areas (e.g. in the mesotrophic Fennoscandian mountains) they represent a substantial part of the crustose lichen flora on rocks, the genus remains poorly represented in herbaria worldwide. We have collected Miriquidica species systematically throughout Norway, representing wide climatic, geographical and ecological gradients, and determined/studied this material through a combinations of different approaches (TLC, microscopy, DNA-sequencing). Our molecular phylogeny of the genus include multiple accessions of all available species as well as extra-Norwegian material. Most Miriquidica species will remain in a monophyletically circumscribed Miriquidica. Some Miriquidica species, however, are clearly extraneous in the genus, and a few species of Lecanora and Lecidea s.lat. belong in Miriquidica. Several new species are described and an extensive amount of phenotypically cryptic species have so far been discovered.
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Haugan, Reidar; Bendiksby, Mika; Rui, Siri & Timdal, Einar
(2013).
Molecular phylogeny and taxonomy of the lichen genus Miriquidica (Ascomycota, Lecanoraceae).
Vis sammendrag
During the years 2010-2012 , the authors participated in the species-project Mapping of Norwegian Lecideoid lichens, funded by the Norwegian Biodiversity Information Centre. Results of the project are presented on the internet (species descriptions, pictures, and distribution data; http://nhm2.uio.no/botanisk/lav/LAVFLORA/LECIDEOID.HTM) and as research articles addressing species-delimitation and phylogeny in selected species groups. In one of these groups, the genus Miriquidica (Hertel & Rambold 1987), which comprises about 30 saxicolous species worldwide, we suspected several unknown species to occur. Many Miriquidica species are difficult to identify without careful chemical (TLC) and microscope investigation. The genus was originally delimited by the presence of miriquidic acid (not all species) and anatomical characters of the apothecia, including asci of the “Lecanora-type”. Although Miriquidica species are not uncommon in alpine areas, and in some areas (e.g. in the mesotrophic Fennoscandian mountains) they represent a substantial part of the crustose lichen flora on rocks, the genus remains poorly represented in herbaria worldwide. During our project, Miriquidica species were systematically collected all over Norway, representing wide climatic, geographical and ecological gradients, and determined/studied through a combinations of different approaches (TLC, microscopy, comparative DNA sequencing). Our molecular phylogeny of the genus include multiple accessions of all available species as well as extra-Norwegian material. Most Miriquidica species will remain in monophyletically circumscribed Miriquidica. Some Miriquidica species, however, were extraneous in the genus, and a few species of Lecanora and Lecidea s.lat. showed to belong in Miriquidica. Several new “phenotypic” species will be described and an extensive amount of phenotypically cryptic species have so far been discovered. The genus is well situated within Lecanoraceae.
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Bendiksby, Mika & Timdal, Einar
(2013).
Kartlegging av lecideoide lav i Norge-2: Oppbygging av et DNA-strekkodebibliotek.
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Bendiksby, Mika & Timdal, Einar
(2013).
Extreme polyphyly and morphological/ecological convergence in the lichen genus Hypocenomyce.
Vis sammendrag
The lichen genus Hypocenomyce consists of ecological specialists growing mainly on burnt wood throughout the boreal zone of the northern hemisphere and also on burnt Eucalyptus in Australia. We have tested hypotheses about five presumed subgroups among the 15 species of Hypocenomyce s.l. (including Pycnora) using a molecular phylogenetic approach. Bayesian, likelihood and parsimony phylogenetic analyses of a dataset consisting of both nuclear (ITS, LSU) and mitochondrial (SSU) ribosomal DNA-regions and a broad Lecanoromycete taxon sampling, mostly support the five presumed subgroups, but two of these were found to be polyphyletic. More surprisingly, our results show that Hypocenomyce s.l. is highly polyphyletic and represents a remarkable example of morphological and ecological convergence. Based on our molecular phylogenetic results, we split Hypocenomyce s.l. into seven genera.
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Bendiksby, Mika; Timdal, Einar; Wolseley, Pat; Thüs, Holger & Vairappan, Charles S.
(2013).
Molecular systematics and taxonomy of Southeast Asian members of the tropical rainforest genus Phyllopsora (Ramalinaceae).
Vis sammendrag
The mainly tropical rainforest genus Phyllopsora (Ramalinaceae) consists of ca. 70 known species. The circumscription of the genus is not clear, and species identification is often difficult due to few morphological characters. The secondary chemistry is rich, however, and offers an important first tool in identification work. We have addressed problems in the current genus and species circumscriptions by molecular methods. About 40 Phyllopsora species have been sequenced so far, and preliminary results from our on-going construction of the genus’ phylogeny and from our taxonomic revision of the species in SE Asia will be presented.
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Haugan, Reidar; Holien, Håkon; Hovind, Annie Beret Ås; Ihlen, Per Gerhard & Timdal, Einar
(2021).
Artsgruppeomtale lav ('Lichenes'). Norsk rødliste for arter 2021.
Artsdatabanken.
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Haugan, Reidar; Holien, Håkon; Hovind, Annie Beret Ås; Ihlen, Per Gerhard & Timdal, Einar
(2021).
Lav ('Lichenes'). Norsk rødliste for arter 2021.
Artsdatabanken.
