Liv Borgen

• Gustafsson, A Lovisa S; Gussarova, Galina; Borgen, Liv; Ikeda, Hajime; Antonelli, Alexandre & Marie-Orleach, Lucas [Vis alle 8 forfattere av denne artikkelen] (2021). Rapid evolution of post-zygotic reproductive isolation is widespread in Arctic plant lineages. Annals of Botany. ISSN 0305-7364. 129(2), s. 171–184. doi: 10.1093/aob/mcab128. Fulltekst i vitenarkiv
• Borgen, Liv (2016). Årets plante, jåblom Parnassia palustris, sett fra et populasjonsgenetisk perspektiv. Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 74(1), s. 7–11.
• Gustafsson, A Lovisa S; Skrede, Inger; Rowe, Heather C.; Gussarova, Galina; Borgen, Liv & Rieseberg, Loren [Vis alle 8 forfattere av denne artikkelen] (2014). Genetics of cryptic speciation within an arctic Mustard, Draba nivalis. PLOS ONE. ISSN 1932-6203. 9(4). doi: 10.1371/journal.pone.0093834. Fulltekst i vitenarkiv
• Marcussen, Thomas & Borgen, Liv (2011). Species delimitation in the Ponto-Caucasian Viola sieheana complex, based on evidence from allozymes, morphology, ploidy levels, and crossing experiments. Plant Systematics and Evolution. ISSN 0378-2697. 291(3-4), s. 183–196. doi: 10.1007/s00606-010-0377-z.
• Borgen, Liv & Guldahl, Ane Senstad (2011). Great-granny's Garden: a living archive and a sensory garden. Biodiversity and Conservation. ISSN 0960-3115. 20(2), s. 441–449. doi: 10.1007/s10531-010-9931-9.
• Bendiksby, Mika; Tribsch, Andreas; Borgen, Liv; Travnicek, Pavel & Brysting, Anne Krag (2011). Allopolyploid origins of the Galeopsis tetraploids - revisiting Müntzing's classical textbook example using molecular tools. New Phytologist. ISSN 0028-646X. 191(4), s. 1150–1167. doi: 10.1111/j.1469-8137.2011.03753.x. Vis sammendrag

  • Whole-genome duplication coupled with hybridization is of prime importance in plant evolution. Here we reinvestigate Müntzing's classical example of allopolyploid speciation; the first report of experimental synthesis of a naturally occurring allopolyploid species, Galeopsis tetrahit. • Various molecular markers (cpDNA, NRPA2, amplified fragment length polymorphisms (AFLPs)) and flow cytometry were surveyed in population samples of subgenus Galeopsis, including two allopolyploid species and their potential diploid parents. • The presence of two divergent copies of single-copy NRPA2 confirms the allopolyploid origins of G. tetrahit and Galeopsis bifida. However, the two allopolyploids do not share the same maternal genome, as originally suggested by Müntzing. The results support independent origins, but not recurrent formation, of the two allotetraploids. Data further indicate frequent gene flow and introgression within ploidy levels, but less so between ploidy levels. • Our results confirm and elaborate on Müntzing's classical conclusion about allopolyploid origins of G. tetrahit and G. bifida. We address questions of general interest within polyploidy research, such as recurrent formation, gene flow and introgression within and between ploidy levels.

• Borgen, Liv (2010). Lobularia, Flora of North America, Volume 7: Magnoliophyta: Salicaceae to Brassicaceae. Oxford University Press. ISSN 9780195318227. s. 597–598.
• Fjellheim, Siri; Jørgensen, Marte Holten; Kjos, Mari & Borgen, Liv (2009). A molecular study of hybridization and homoploid hybrid speciation in Argyranthemum (Asteraceae) on Tenerife, the Canary Islands. Botanical journal of the Linnean Society. ISSN 0024-4074. 159(1), s. 19–31. Vis sammendrag

  Examples of recurrent homoploid hybrid speciation are few. One often-cited example is Argyranthemum sundingii. This example includes two described species, A. lemsii and A. sundingii, resulting from reciprocal hybridization between A. broussonetii and A. frutescens on Tenerife. The four species and artificial F-1 and F-2 hybrids have previously been investigated morphologically and cytologically. Here, we examine population differentiation based on amplified fragment length polymorphism to get a better understanding of the genetic relationships among the species and the extent of hybridization. We aim to investigate if there is molecular support for treating the hybrid species as one taxon. Seven parental and four hybrid species populations (149 individuals) were analysed and we scored 85 polymorphic markers. A few (2-5) were private to each species but variably present and mostly rare. Our principal coordinate, STRUCTURE and BAPS analyses and AMOVA resulted in a clear separation of the parental species. The hybrid species were genetically less divergent but not identical. Our data indicate that hybridization and introgression are common in all these species on Tenerife and support the hypothesis that homoploid hybrid speciation has occurred repeatedly. Intrinsic post-zygotic barriers are notoriously weak in Argyranthemum and reproductive isolation and speciation result primarily from strong ecological selection. (C) 2009 The Linnean Society of London, Botanical Journal of the Linnean Society, 2009, 159, 19-31.

• Fjellheim, Siri; Jørgensen, Marte Holten; Kjos, Mari & Borgen, Liv (2009). A molecular study of hybridization and homoploid hybrid speciation in Argyranthemum (Asteraceae) on Tenerife, the Canary Islands. Botanical journal of the Linnean Society. ISSN 0024-4074. 159(1), s. 19–31. doi: 10.1111/j.1095-8339.2008.00947.x.
• Skrede, Inger; Borgen, Liv & Brochmann, Christian (2009). Genetic structuring in three closely related circumpolar plant species: AFLP versus microsatellite markers and high-arctic versus arctic-alpine distributions. Heredity. ISSN 0018-067X. 102(3), s. 293–302. doi: 10.1038/hdy.2008.120.
• Skrede, Inger; Brochmann, Christian; Borgen, Liv & Rieseberg, LH (2008). Genetics of intrinsic postzygotic isolation in a circumpolar plant species, Draba Nivalis (Brassicaceae). Evolution. ISSN 0014-3820. 62, s. 1840–1851. doi: 10.1111/j.1558-5646.2008.00418.x.
• Borgen, Liv (2007). Etnobotanikk. I Jørgensen, Per M. (Red.), Botanikkens historie i Norge. Fagbokforlaget. ISSN 978-82-450-0499-1. s. 116–123.
• Borgen, Liv & Jørgensen, Per Magnus (2007). Schübeler, Frederik Christian. I Jørgensen, Per M. (Red.), Botanikkens historie i Norge. Fagbokforlaget. ISSN 978-82-450-0499-1. s. 349–351.
• Grundt, Hanne Hegre; Kjølner, Siri; Borgen, Liv; Rieseberg, LH & Brochmann, Christian (2006). High biological species diversity in the arctic flora. Proceedings of the National Academy of Sciences of the United States of America. ISSN 0027-8424. 103, s. 972–975.
• Marcussen, Thomas; Borgen, Liv & Nordal, Inger (2005). New distributional and molecular information call into question the systematic position of the West Asian Viola sintenisii (Violaceae). Botanical journal of the Linnean Society. ISSN 0024-4074. 147, s. 91–98.
• Guldahl, AS; Gabrielsen, Tove M; Scheen, Anne-Cathrine; Borgen, Liv; Steen, SW & Spjelkavik, S [Vis alle 7 forfattere av denne artikkelen] (2005). The Saxifraga rivularis complex in Svalbard: Molecules, ploidy and morphology. Flora: Morphologie, Geobotanik, Oekophysiologie. ISSN 0367-2530. 200, s. 207–221.
• Grundt, Hanne H.; Obermayer, Renate & Borgen, Liv (2005). Ploidal levels in the arctic-alpine polyploid Draba lactea (Brassicaceae) and its low-ploid relatives. Botanical journal of the Linnean Society. ISSN 0024-4074. 147, s. 333–347.
• Brochmann, Christian; Brysting, Anne Krag; Alsos, Inger Greve; Borgen, Liv; Grundt, Hanne Hegre & Scheen, Anne-Cathrine [Vis alle 7 forfattere av denne artikkelen] (2004). Polyploidy in arctic plants. Biological Journal of the Linnean Society. ISSN 0024-4066. 82, s. 521–536.
• Borgen, Liv (2003). Genome organization in diploid hybrid species of Argyranthemum (Asteraceae) in the Canary Islands. Botanical journal of the Linnean Society. ISSN 0024-4074. 141, s. 491–501.
• Borgen, Liv (2003). Parnassia palustris: a genetically diverse species in Scandinavia. Botanical journal of the Linnean Society. ISSN 0024-4074. 142, s. 347–372.
• Hagen, Aslaug Ragnhild; Borgen, Liv; Elven, Reidar; Stabbetorp, Odd Egil & Brochmann, Christian (2002). The arctic-alpine polyploids Cerastium alpinum and C. nigrescens (Caryophyllaceae) in a sympatric situation: breakdown of species integrity? Plant Systematics and Evolution. ISSN 0378-2697. 230, s. 203–219.
• Borgen, Liv (2002). Clypeola maritima L., Sp. Pl.:62, 1 May 1753 = Lobularia maritima (L.) Desv. P.533 in: Cafferty, S. & Jarvis, C. E. (eds), Typification of Linnaean plant names in Brassicaceae (Cruciferae). Taxon. ISSN 0040-0262. 51, s. 529–537.
• Borgen, Liv & Often, Anders (2001). Stellaria 11-15. I Jonsell, Bengt & Karlsson, T. (Red.), Flora Nordica 2. The Bergius Foundation, The Royal Swedish Academy of Sciences, Stockholm.. s. 129–133.
• Grundt, Hanne Hegre; Borgen, Liv & Elven, Reidar (2001). Infraspecific variation in Cerastium alpinum s. lat. (Caryophyllaceae) in Central Norway. Nordic Journal of Botany (NJB). ISSN 0107-055X. 20(6), s. 641–653.
• Marcussen, Thomas; Borgen, Liv & Nordal, Inger (2001). Viola hirta (Violaceae) and its relatives in Norway. Nordic Journal of Botany (NJB). ISSN 0107-055X. 21(1), s. 5–17. Vis sammendrag

  Viola hirta is widespread in most of Europe but rare in Norway, occurring in four isolated enclaves around the Oslo Fjord. Allozymic and morphological patterns of variation were studied in 320 individuals of Viola hirta (2n = 20) and its relatives V. collina (2n = 20), V. odorata (2n = 20), and V. suavis (2n = 40) from 22 Norwegian populations. Ongoing introgression from both V. collina and V. odorata, as hypothesised by several authors, was not supported by our data. All hybrids proved to be primary ones and, apparently, introgression is rare. However, allozyme markers suggested that V. hirta may have been affected by ancient introgression from V. collina and possibly also from the East European V. ambigua (2n = 40) during its evolutionary history. Geographical differentiation, presumably relatively recent and post-glacial, was evident in V. hirta. Variation occurred also between and within V. odorata populations, most likely a consequence of independent introductions by man. No variation was observed in V. collina or in V. suavis. A key to the Norwegian species of subsection Viola and their primary hybrids based on morphological characters is provided in Flora Nordica note No. 26.

• Guldahl, A. S.; Borgen, Liv & Nordal, Inger (2001). Variation in the Festuca brachyphylla (Poaceae) complex in Svalbard, elucidated by chromosome numbers and isozymes. Botanical journal of the Linnean Society. ISSN 0024-4074. 137, s. 107–126.
• Borgen, Liv (2000). Professor Schübelers plass i norsk hagebrukshistorie. I Moe, D.; Salvesen, P.H. & Øvstedal, D.O. (Red.), Historiske hager. Fagbokforlaget. s. 22–27.
• Marcussen, Thomas & Borgen, Liv (2000). Allozymic variation and relationships within Viola subsection Viola (Violaceae). Plant Systematics and Evolution. ISSN 0378-2697. 223, s. 29–57. Vis sammendrag

  Allozyme markers from ten European taxa of Viola subsection Viola suggests that this group is allotetraploid, based on x=5. All taxa had distinct multilocus phenotypes except V. alba sspp. alba and scotophylla, which were identical and different from ssp. dehnhardtii. Variation was consistently higher in Mediterranean populations than in North European ones. Hybridisation seems extensive but putative F1 hybrids were distinctly less fertile than the parental species. Nevertheless, increased fertility in later-generation hybrids and shared band patterns among taxa indicate an important role of hybridisation and introgression in past and present evolution within the subsection. The octoploid V. ambigua shows affinity to V. hirta (tetraploid). The octoploid V. suavis probably originated from V. pyrenaica and other unidentified tetraploids, and high variability suggests polytopy or even polyphyly. The stoloniferous condition (series Flagellatae) seems to be primitive in the subsection but the reduction of stolons (series Eflagellatae) may have originated multiple times.

• Brochmann, Christian; Borgen, Liv & Stabbetorp, Odd Egil (2000). Multiple diploid hybrid speciation of the Canary Island endemic Argyranthermum sundingii (Asteraceae). Plant Systematics and Evolution. ISSN 0378-2697. 220, s. 77–92.
• Brysting, Anne Krag & Borgen, Liv (2000). Isozyme analysis of the Cerastium alpinum-C. arcticum complex (Caryophyllaceae) supports a splitting of C. arcticum Lange. Plant Systematics and Evolution. ISSN 0378-2697. 220, s. 199–221.
• Borgen, Liv; Berg, Rolf Y.; Sjølie, H.; Torvik, S.E. & Vejrup, K. (2000). Mating system variation in three species of flowering plants on Hovedøya, Oslo, Norway. Avhandlinger (Det Norske Videnskaps-Akademi). 1. Matematisk-Naturvitenskapelig Klasse. ISSN 0549-7019. 39, s. 67–74.
• Borgen, Liv (1999). Genetic variation in Minuartia (Caryophyllaceae) in Svalbard. Nordic Journal of Botany (NJB). ISSN 0107-055X. 19, s. 179–192.
• Grundt, Hanne Hegre; Borgen, Liv & Elven, Reidar (1999). Aspects of reproduction in Cerastium alpinum on calcic and ultramafic soils in Central Norway. Nordic Journal of Botany (NJB). ISSN 0107-055X. 19, s. 447–453.

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• Borgen, Liv (2014). Botanisk hage 1814-2014. Historien om en hage. Forlaget Press. ISBN 9788275477048. 335 s.
• Bjorå, Charlotte Sletten; Bjureke, Kristina; Borgen, Liv; Danielsen, Dag Inge; Ervik, Finn & Finnanger, Anne [Vis alle 8 forfattere av denne artikkelen] (2011). Botanisk hage. En oase i Oslo. Universitetet i Oslo. ISBN 978-82-79-70025-8. 127 s.
• Borgen, Liv (2008). Fjellhagen - Botanisk hages vakreste anlegg. Universitetet i Oslo. ISBN 978-82-79-70008-1. 96 s.
• Borgen, Liv (2006). Urtegleder - Krydderplanter i Urtehagen i Botanisk hage. Universitetet i Oslo. ISBN 978-82-7970-007-4. 128 s.
• Borgen, Liv (2004). Schübeler, Frederik Christian ("Fritz"), 1815-92, botaniker [biografi]. Kunnskapsforlaget. 2 s.

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• Haraldsen, Kirsten Borse; Pedersen, Oddvar; Borgen, Liv & Nordal, Inger (2019). Rolf Y. Berg – bibliografi . Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 77(1), s. 30–34.
• Borgen, Liv; Brochmann, Christian; Haraldsen, Kirsten Borse; Nordal, Inger & Pedersen, Oddvar (2019). Rolf Yngvar Berg 1925–2018 . Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 77(1), s. 24–30.
• Nordal, Inger; Borgen, Liv; Haraldsen, Kirsten Borse & Pedersen, Oddvar (2019). Et «dypdykk» i et livslangt forsknings-engasjement: Rolf Y. Berg . Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 77(1), s. 35–43.
• Høiland, Klaus & Borgen, Liv (2018). Professor Frederik Christian Schübeler som feltbotaniker. Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 76(1), s. 47–62.
• Borgen, Liv (2017). Frederik Christian Schübelers virke i Botanisk hage på Tøyen under mottoet "Dyrk verdifuldere vekster". Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 75, s. 91–112.
• Borgen, Liv (2015). Intervju om veksthusene i Botanisk hage. [Radio]. NRK P2 "Ekko" - intervjuet fant sted i Botanisk hage.
• Borgen, Liv (2015). Botanisk hages historie.
• Østhagen, Haavard & Borgen, Liv (2015). Årsmøte DnR i Stavanger 22.-23. mai 2015. Lapprosen. ISSN 1504-6702. 18(3), s. 26–27.
• Borgen, Liv (2015). Bokanmeldelse: Praktverk om norske klosterplanter og norske middelalderklostre. Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 73(253).
• Borgen, Liv (2015). Botanisk hage på Tøyen og Frederik Christian Schübelers betydning for norsk hagebruk. Lokalhistorisk magasin. ISSN 0802-8931. s. 4–9.
• Borgen, Liv & Johansen, Bjørn Vidar (2014). Botanisk hage 1814-2014. Historien om en hage, Botanisk hage 1814-2014. Historien om en hage.. Forlaget Press. ISSN 9788275477048.
• Borgen, Liv (2014). Botanisk hage 1814-2014. St. Hallvard. ISSN 0036-2859. 2, s. 4–21.
• Borgen, Liv (2014). Botanisk hage grunnlagt i 1814: Viktig for den unge nasjonen. Apollon : Forskningsmagasin for Universitetet i Oslo. ISSN 0803-6926. 2, s. 18–21.
• Bendiksby, Mika; Tribsch, Andreas; Borgen, Liv; Travnicek, Pavel & Brysting, Anne Krag (2012). Revisiting Müntzing's classical textbook example of allotetraploid Galeopsis tetrahit - using molecular tools.
• Gustafsson, A Lovisa S; Borgen, Liv; Gusarova, Galina & Brochmann, Christian (2011). Cryptic Speciation in Arctic Plants.
• Gustafsson, A Lovisa S; Brochmann, Christian; Gusarova, Galina & Borgen, Liv (2011). Cryptic Speciation in the Arctic.
• Gustafsson, A Lovisa S; Parisod, Christian; Schranz, Eric; Borgen, Liv; Gusarova, Galina & Brochmann, Christian (2010). Genetics of Intrinsic Postzygotic Isolation in Draba nivalis (Brassicaceae).
• Gustafsson, A Lovisa S; Brochmann, Christian; Borgen, Liv & Gusarova, Galina (2010). High Biological Species Diversity in the Arctic Flora?
• Borgen, Liv (2009). Hva er et gresskar? Botanikk, systematikk og historikk. I Ulfeng, A & Ulfeng, I J (Red.), Gresskar - mye mer enn Halloween - matglede og botanisk eventyr. Møklegaards Trykkeri AS. ISSN 978-82-303-1262-9. s. 13–27.
• Borgen, Liv (2009). Einer - den nyttigste planten i norsk natur? Norsk institutt for skog og landskap.
• Gustafsson, Lovisa; Borgen, Liv; Gusarova, Galina & Brochmann, Christian (2009). High Biological Species Diversity in the Arctic Flora?
• Borgen, Liv & Guldahl, Ane Senstad (2009). Great-Granny’s Garden – A Living Archive and a Sensory Garden.
• Borgen, Liv (2008). Verifisering av hasjplante. [Avis]. VG.
• Skrede, Inger; Brochmann, Christian; Borgen, Liv & Rieseberg, Loren h (2008). Sibling speciation in the diploid, selfing Draba nivalis (Brassicaceae).
• Skrede, Inger; Borgen, Liv; Brochmann, Christian & Rieseberg, Loren (2007). Sibling speciation in arctic diploid Draba (Brassicaceae).
• Borgen, Liv; Fjellheim, S & Kjos, Mari (2007). AFLP data confirm recurrent homoploid hybrid speciation in Argyranthemum (Asteraceae).
• Skrede, Inger; Borgen, Liv; Brochmann, Christian & Rieseberg, Loren (2007). Sibling speciation in arctic diploid Draba (Brassicaceae). Vis sammendrag

  Incomplete to fully developed postzygotic sterility barriers within species were recently demonstrated in three circumpolar species of the genus Draba, suggesting numerous sibling species (Grundt et al., 2006; PNAS, 103, 972-975). Because the three species are selfers and of recent, Pleistocene origin, Grundt et al. argued that hybrid sterility evolved quickly and possibly by genetic drift. The present study employs genetic mapping and quantitative trait locus (QTL) analyses to determine whether the sterility has a chromosomal or genic basis and, if the latter, whether the sterility is caused by interactions among alleles at a single locus or by interactions among alleles at different loci. A F2 population of 383 individuals was generated by crossing two Draba nivalis plants, one from Alaska and one from Norway (F1 hybrids were ~30 % fertile). All F2 plants have been tested for pollen viability, seed set, and growth rate, and were on average 62 % fertile for the two fertility measures. A microsatellite-enriched library has been made for Draba, and 50 loci have been genotyped for all individuals. QTL analyses identified five loci underlying seed fertility and two affecting reduced pollen viability. Seed fertility QTLs were weakly underdominant, an observation most consistent with single locus sterility, such as that caused by chromosomal rearrangements. In contrast, pollen sterility QTLs exhibited additive inheritance patterns as might be expected for Dobzhansky-Muller incompatibilities. The small number of sterility QTLs is consistent with results from other plant groups, which imply that fewer loci contribute to hybrid sterility in plants than in Drosophila.

• Borgen, Liv (2006). Ex situ/ conservation and related activities in the Botanical Garden, Natural History Museum, University of Oslo.
• Skrede, Inger; Borgen, Liv; Brochmann, Christian & Rieseberg, Loren (2006). The mechanisms of postzygotic reproductive isolation in Draba nivalis (Brassicaceae). Vis sammendrag

  Incomplete to fully developed postzygotic sterility barriers within species were recently demonstrated in three circumpolar species of the genus Draba (Grundt et al., 2006; PNAS, 103, 972-975). Because the three species are selfers and of recent, Pleistocene origin, Grundt et al. argued that hybrid sterility evolved quickly and possibly by genetic drift. The present study employs a combination of QTL and meiotic analyses to determine whether the sterility has a chromosomal or genic basis and, if the latter, whether the sterility is caused by interactions among alleles at a single locus or by interactions among alleles at different loci. We have generated an F2 population of ~400 individuals by crossing Draba nivalis plants from Alaska and Norway (F1 hybrids were ~50% fertile). All F2 plants have been tested for pollen viability, seed set, and biomass. To genotype the F2 population, a microsatellite-enriched library has been made for Draba, and 60 microsatellites have been developed. So far, 23 markers have been analyzed for all individuals.

• Borgen, Liv (2006). Presentasjon av urtehagen i Botanisk hage.
• Borgen, Liv (2005). Plantearven - vårt felles ansvar. Palmebladet. ISSN 1503-4593. 6, s. 6–8.
• Borgen, Liv; Jørgensen, Marte Holten & Obermayer, Renate (2005). Why is it so difficult to obtain high quality AFLP profiles in Argyranthemum?
• Obermayer, Renate; Grundt, Hanne Hegre & Borgen, Liv (2005). Ploidy patterns in the arctic-alpine polyploid Draba lactea Adams (Brassicaceae) and its low-ploid relatives.
• Borgen, Liv; Jørgensen, Marte & Obermayer, Renate (2005). Why is it so difficult to obtain high quality AFLP profiles in Argyranthemum?
• Borgen, Liv (2004). Planter til nytte og glede, død og fordervelse.
• Borgen, Liv (2004). Medisinplanter i norsk natur.
• Borgen, Liv (2004). Historien om et levende fossil, urtidstreet Metasequoia glyptostroboides. Blyttia : Norsk botanisk forenings tidsskrift. ISSN 0006-5269. 62, s. 63–66.
• Borgen, Liv (2004). Fjellhagen - noen av høydepunktene i sommerhalvåret. Palmebladet. ISSN 1503-4593. 3, s. 3–5.
• Borgen, Liv (2004). Botanisk hages bestyrer 1866-1892, Frederik Christian Schübeler. Palmebladet. ISSN 1503-4593. 4, s. 2–3.
• Borgen, Liv (2004). Rapport fra en studiereise. Palmebladet. ISSN 1503-4593. 4, s. 21–23.
• Obermayer, Renate; Grundt, Hanne Hegre & Borgen, Liv (2004). Monitoring ploidy patterns in arctic-alpine polyploid Draba lactea Adams (Brassicaceae) and its low-ploid relatives.
• Borgen, Liv (2003). Conservation of important horticultural plants in European Botanic Gardens: a case study from Norway. Scripta Botanica Belgica. ISSN 0779-2387.
• Borgen, Liv (2003). Februarblomstring i Botanisk hage i Oslo. Norsk hagetidend : populært tidsskrift for hagedyrkere. ISSN 0029-1986.
• Borgen, Liv (2003). Gresskar og Halloween i Botanisk hage - mangfold og glede, grøss og gru. Palmebladet. ISSN 1503-4593. 1, s. 9–10.
• Borgen, Liv (2003). Gresskar. Palmebladet. ISSN 1503-4593. 1, s. 11–12.
• Borgen, Liv (2003). Fjellhagen. Palmebladet. ISSN 1503-4593. 2, s. 3–6.
• Brochmann, Christian; Brysting, Anne Krag; Grundt, Hanne Hegre; Borgen, Liv & Elven, Reidar (2003). Polyploidy in arctic plants.
• Borgen, Liv (2002). Vår i Kew Gardens.
• Grundt, Hanne Hegre; Popp, M.; Borgen, Liv; Elven, Reidar; Brochmann, Christian & Oxelman, B. (2002). The history of the circumpolar allopolyploid Draba lactea based on homoeologous nuclear DNA sequences.
• Borgen, Liv (2002). Bevaring av plantegenetiske ressurser i klonarkiv.
• Borgen, Liv (2002). Parnassia palustris: a genetically diverse species in Scandinavia.
• Borgen, Liv (2001). Fra naturens verden. [Radio]. Oslo. : Radioens P1..
• Borgen, Liv (2000). Legeplanter i norsk flora. I. Våre nyttevekster. ISSN 0332-7280. 95, s. 54–61.
• Borgen, Liv (2000). Legeplanter i norsk flora. II. Våre nyttevekster. ISSN 0332-7280. 95, s. 92–104.
• Borgen, Liv (2000). Purpursolhatt, Echinacea purpurea. Botaniske Ark. 3.
• Borgen, Liv (2000). Eseltistel, Onopordon acanthium. Botaniske Ark. 4.
• Borgen, Liv (2000). Dufthagen på Tøyen. Guidehefte til Botanisk hages "Dufthage". Botanisk hage og museum..
• Borgen, Liv (2000). The Rock Garden. Highlights. Selected atractions Natural History Museums and Botanical Garden.. s. 8–17.
• Borgen, Liv (1999). Glimt fra humlens og humlehagenes historie i Norge. Våre nyttevekster. ISSN 0332-7280. 94, s. 45–56.
• Scheen, Anne-Cathrine; Brochmann, Christian; Spjelkavik, S.; Borgen, Liv & Elven, Reidar (1999). The Draba lactea/fladnizensis complex (Brassicaceae) in Svalbard: relationships and evolution within the group.
• Bjureke, Kristina; Borgen, Liv; Finnanger, Anne & Skinnarland, Nils Ifver (2002). Gresskar - mangfold og glede, grøss og gru. Universitetets naturhistoriske museer og botaniske hage.
• Borgen, Liv (2000). Kompendium i enzymelektroforese. Lab.-manual for BB 300, Botanikk, innføringskurs til hovedfag. [Oppdatert versjon.]. Universitetet i Oslo.

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